July 9, 1900 Born in Landeshut Silesia, Germany (now Poland)
1918-1919 University of Breslau (Germany)
1919-1920 University of Heidelberg (Germany)
1920-1921 University of Freiburg (Germany)
1921-1922 University of München (Germany)
1922-1925 University of Freiburg (Germany)
1925 Ph.D. in Zoology (Experimental Embryology) under Professor H. Spemann, Nobel Laureate. University of Freiburg
1925-1926 Research Associate, University of Göttingen (Prof. A. Kühn)
1926-1927 Research Associate, Kaiser Wilhelm Institute for Biology (Prof. O. Mangold) Berlin - Dahlem, Germany
1927-1932 Privatdozent, Instructor, Zoological Institute, University of Freiburg
1932-1935 Rockefeller Fellow, Instructor, Department of Zoology, University of Chicago
1935-1939 Assistant Professor of Zoology, Washington University, St. Louis
1939-1941 Associate Professor of Zoology, Washington University, St. Louis
1941-1968 Professor of Zoology, Washington University, St. Louis
1968-1969 Edward Mallinckrodt Distinguished Service Professor of Biology, Washington University, St. Louis
1969- Edward Mallinckrodt Distinguished Service Professor Emeritus of Biology, Washington University, St. Louis
1941-1966 Chairman, Department of Zoology, Washington University, St. Louis
1936-1945 Instructor, later Director, Embryology Course, Marine Biology Laboratory, Woods Hole, MA
1947 (Spring) Special Research Fellow, National Institutes of Health, Bethesda, MD
1947 (Summer) Visiting Professor, Zoology Department, University of Chicago
1950 (Spring) Visiting Professor, Department of Biology, Massachusetts Institute of Technology
1950 (Summer) Visiting Professor, Department of Zoology, University of California, Berkeley
June 12, 2001 Died, St. Louis, Missouri


Member American Society of Zoologists (President, 1955)
American Association of Anatomists
American Association for the Advancement of Science (AAAS) (Vice-President, Chairman, Section F, 1960)
American Society for Developmental Biology (President, 1950, 1951)
Society for Developmental Biology
International Society for Cell Biology
International Society of Developmental Biologists
Honorary
Member International Society for Developmental Neuroscience

Honors National Academy of Science (U.S.A.), 1953
American Academy of Arts and Sciences, 1959
Sigma XI
Phi Beta Kappa
Honorary Doctorate of Science, Washington University, May 21, 1976
F.O. Schmitt Medal and Prize in Neuroscience, Oct. 20, 1976
Wakeman Award in Neuroscience, 1978
Ross G. Harrison - ISDB Prize in Developmental Biology of the International Society of Developmental Biologists, Aug. 21, 1981
Louisa Gross Horwitz Prize in Cell Biology and Developmental NeuroBiology, Columbia University, Oct. 17, 1983
Honorary Doctorate of Philosophy, University of Uppsala, Sweden, 1984
Ralph Gerard Prize and Medal of the Society for Neuroscience, 1985
Fidia-Georgetown Award in Neuroscience, May, 1987
National Medal of Science, Oct. 18, 1989
Karl Spencer Lashley Award of the American Philosophical Society, April, 1990
Honorary Member, American Society of Zoologists, 1990
Honorary Doctorate, Rockefeller University, 1999
(First) Lifetime Achievement Award, Society for Developmental Biology, 2000.